The lethal period of development starts at the first larval instar and lasts the whole period of development, with peaks at the first and second larval instar stages. Pupariation is delayed for 60-80 hours and is abnormal; some animals do not show pupal moulting and die at the stage of gas-bubble formation, others die at the stage of eye pigmentation and bristle formation. Those surviving to imago formation generally die before or at the moment of eclosion, while the 2-5% that manage to eclose live for only a few days.
Fertility is unaffected.