groC105/groE28 mutants are semi-viable, escapers have rough eyes and thickened wing vein L5.
Heterozygotes that are also heterozygous for groBFP2 have eye defects that are similar but not identical to groBFP2 homozygotes. Homozygous mutant clones in the eye show ommatidial defects, often having extra photoreceptors.
Weak neurogenic phenotype.